Magnificent bryozoan	Status LU: established. 1st record: ~2011.
	Schwamp-Moosdéierchen	Status Eur.: established.
	Pectinatelle	RA: ISEIA: C1. Harmonia+: n/a.
	Schwammartiges Moostierchen	Wikipedia: | Wikispecies:
	n/a	Back to the list of invertebrates

Brief description

Pectinatella magnifica, the magnificent bryozoan, is a member of the Bryozoa phylum, in the order Plumatellida. It is a colony of organisms that bind together; these colonies can sometimes be 60 centimeters in diameter. These organisms can be found mostly in North America with some in Europe. They are often found attached to objects, but can be found free floating as well. They form a translucent body with many star-like blooms along the outside. The density of the organism is similar to that of gelatin, and is easily breakable into smaller chunks (Wikipedia contributors 2018).

Status and distribution in Luxembourg

In 2012 numerous colonies of Pectinatella magnifica (Leidy, 1851) (Bryozoa, Phylactolaemata) were discovered in the reservoir of Esch-sur-Sûre (Luxembourg) fed by the river Sûre. The colonies were particularly abundant in the shallow, warm and nutrient-rich water near the riverbank, but some colonies were spotted by divers in the reservoir at a depth of 8-9 m in one site and more than 20 m in another site. There is reliable evidence that Pectinatella was present, but less conspicuous and as such not identified, in 2011 and possibly already in 2010. P. magnifica was hitherto unrecorded in Luxembourg, but known from a site near the German-Luxembourg border near Nennig (Germany, Saarland) where statoblasts were found in 2001 (Massard et al. 2013).

Risk assessment

ISEIA protocol

C1 (3+2+1+1) (Ries et al. 2017: 68).

Harmonia+ protocol

Not assessed yet.

Worldwide distribution

Bibliography

• GBIF, 2019. Pectinatella magnifica (Leidy, 1851) in GBIF Secretariat (2019). GBIF Backbone Taxonomy. Checklist dataset https://doi.org/10.15468/39omei [accessed 2020-03-13]
• Massard, J.A., G. Geimer & E. Wille, 2013. Apparition de Pectinella magnifica (Leidy, 1851) (Bryozoa, Phylactolaemata) dans le lac de barrage d’Esch-sur-Sûre (Luxembourg). Bulletin de la Société des naturalistes luxembourgeois 114: 131-148.
• Massard, J.A. & G. Geimer, 2015. L’’histoire de la recherche bryozoologique au Luxembourg (Phylactolémates et Gymnolémates d’eau douce). Bull. Soc. Nat. luxemb. 116: 373-379. [PDF 1,35 MB]
• MNHNL, iNaturalist & GBIF, 2021. MNHNL-mdata, online portal combining species observation from Recorder-Lux, iNaturalist and GBIF. National Museum of Natural History, Luxembourg. URL: https://mdata.mnhn.lu [Accessed 2021-04-14]
• Ries, C., A. Arendt, C. Braunert, S. Christian, A. Dohet, A. Frantz, G. Geimer, M. Hellers, J. A. Massard, X. Mestdagh, R. Proess, N. Schneider & M. Pfeiffenschneider, 2017. Environmental impact assessment and black, watch and alert list classification after the ISEIA Protocol of invertebrates in Luxembourg. Bull. Soc. Nat. luxemb. 119: 63-70. [PDF 360 KB]
• Wikipedia contributors, 2018. ‘Pectinatella magnifica’, Wikipedia, The Free Encyclopedia, 3 December 2018, 23:25 UTC, <https://en.wikipedia.org/w/index.php?title=Pectinatella_magnifica&oldid=871869936> [accessed 2020-03-13]

 Page content last updated on 2023-08-21. Last proofread by Caroline Grounds on 2019-11-12.

	Varroa mite	Status LU: established. 1st record: 1985.
	Varroamilb	Status Eur.: established.
	n/a	RA: ISEIA: A3, Black List. Harmonia+: n/a.
	Varroamilbe	Wikipedia: | Wikispecies: | CABI
	Varroamijt	Back to the list of invertebrates

Brief description

Varroa destructor Anderson & Trueman, 2000 is an external parasitic mite that attack and feeds on the honey bees Apis cerana and Apis mellifera. The disease caused by the mites is called varroosis. The Varroa mite can only reproduce in a honey bee colony. It attaches to the body of the bee and weakens the bee by sucking fat bodies. The species is a vector for at least five debilitating bee viruses, including RNA viruses such as the deformed wing virus (DWV). A significant mite infestation leads to the death of a honey bee colony, usually in the late autumn through early spring. The Varroa mite is the parasite with possibly the most pronounced economic impact on the beekeeping industry. Varroa is considered to be one of multiple stress factors contributing to the higher levels of bee losses around the world.

The adult female mite is reddish-brown in color, while the male is white. Varroa mites are flat, having a button shape. They are 1–1.8 mm long and 1.5–2 mm wide, and have eight legs. Mites reproduce on a 10-day cycle (Wikipedia contributors 2020).

Status and distribution in Luxembourg

Varroa destructor Anderson & Trueman, 2000 was observed for the first time in Luxembourg in 1985 (Meisch 1986). Claude Meisch recalls: “At that time it was a delicate matter, no beekeeper wanted to admit that he had the parasite in his hives. I remember well that I was the first to admit in meetings that I had found Varroa in my hives. Probably Varroa was already widespread in the whole country at that time” (Meisch 2018).

In the 1980s, Varroa spread throughout the country and is nowadays present in every beehive.

There is no distribution map available because no data has been entered into the Recorder-Lux database so far (MNHNL, iNaturalist & GBIF 2019).

Risk assessment

ISEIA protocol

A3 (3+3+3+3) = Black List (Ries et al. 2017: 68).

Harmonia+ protocol

Not assessed yet.

Worldwide distribution

Bibliography

• CABI, 2008. Varroa destructor Anderson & Trueman, 2000 [original text by Claire Beverley]. In: Invasive Species Compendium. Wallingford, UK: CAB International. URL: www.cabi.org/isc [accessed 2019-11-26]
• MNHNL, iNaturalist & GBIF, 2019. MNHNL-mdata, online portal combining species observation from Recorder-Lux, iNaturalist and GBIF. National Museum of Natural History, Luxembourg. URL: https://mdata.mnhn.lu [Accessed 2019-10-24]
• Meisch, C., 1986. Die Varroamilbe. Geschichte der Ausbreitung, Portrait und Biologie. Pp. 174-177 in: Livre d’or du centenaire 1886-1986. Fédération des unions d’apiculteurs du grand-duché de Luxembourg.
• Meisch, C., 2018. Personal communication to Lucie Lentz during Summer 2018.
• Ries, C., A. Arendt, C. Braunert, S. Christian, A. Dohet, A. Frantz, G. Geimer, M. Hellers, J. A. Massard, X. Mestdagh, R. Proess, N. Schneider & M. Pfeiffenschneider, 2017. Environmental impact assessment and black, watch and alert list classification after the ISEIA Protocol of invertebrates in Luxembourg. Bull. Soc. Nat. luxemb. 119: 63-70. [PDF 360 KB]
• Wikipedia contributors, 2020. ‘Varroa destructor’, Wikipedia, The Free Encyclopedia, 19 February 2020, 15:24 UTC, <https://en.wikipedia.org/w/index.php?title=Varroa_destructor&oldid=941598994> [accessed 2020-03-13]

 Page content last updated on 2021-05-14. Last proofread by Caroline Grounds on 2019-12-12.

	n/a	Status LU: established. 1st record: 2009.
	Roude Schneider	Status Eur.: established.
	n/a	RA: ISEIA: B3, Watch List. Harmonia+: n/a.
	Apenninen-Kanker	Wikipedia:   | Wikispecies: n/a (2020) | CABI
	Rode hooiwagen	Back to the list of invertebrates

Brief description

Opilio canestrinii is a species of harvestman. Males reach a body length up to 6 mm, females up to 8 mm. While males are yellowish brown to reddish, females are lighter. Males have dark legs, but yellow coxae and “knees”; the legs of females show alternately light and dark rings. The backs of females sport a dark, saddle-like pattern with a light longitudinal stripe in the middle. Adults can be found from June to December.

O. canestrinii probably originates from Italy, but has invaded Central Europe since the late 1970, and has since almost everywhere replaced the similar O. parietinus. It is most often found on house walls (Wikipedia contributors 2019).

Status and distribution in Luxembourg

The first documented occurrence of the species in Luxembourg dates from 2009. It was found by C. Muster on 20th August 2009 in Fort Berlaimont (Municipality of Luxembourg).

Opilio canestrinii is the second most common harvestman in Luxembourg and can be found in every region of the country.

Opilio canestrinii colonises a number of open habitats independently of their moisture levels. It is very successful in man-made landscapes, even in bigger cities. It is a strong competitor to native species and seems able to displace or even replace them. There are indicators that the similar Opilio parietinus, traditionally a common species on house walls, has nearly vanished, just a few years after the arrival of O. canestrinii.

Several sources describe this harvestman as a typical species of urban areas. In Luxembourg however, the data provided from different projects show evidence that the species occurs in great numbers and important percentages even in close-to-nature areas. Since the 1960s, O. canestrinii has spread extremely quickly northwards through Europe and has become a more or less common species in their new areas.

Risk assessment

ISEIA protocol

B3 (3+2+3+1) = Watch List (Ries et al. 2017: 68).

Harmonia+ protocol

Not assessed yet.

Worldwide distribution

Bibliography

• CABI, 2019. Opilio canestrinii. In: Invasive Species Compendium. Wallingford, UK: CAB International. URL: www.cabi.org/isc [accessed 2020-03-05]
• GBIF, 2020. Opilio canestrinii in GBIF Secretariat (2019). GBIF Backbone Taxonomy. Checklist dataset https://doi.org/10.15468/39omei [accessed 2020-03-05]
• MNHNL, iNaturalist & GBIF, 2020. Opilio canestrinii in MNHNL-mdata, online portal combining species observation from Recorder-Lux, iNaturalist and GBIF. National Museum of Natural History, Luxembourg. URL: https://mdata.mnhn.lu [Accessed 2020-03-05]
• Muster, C. & M. Meyer, 2014. Verbreitungsatlas der Weberknechte des Großherzogtums Luxemburg. Ferrantia 70. Musée national d’histoire naturelle, Luxembourg, 112 S. (PDF 3.8 MB)
• Ries, C., A. Arendt, C. Braunert, S. Christian, A. Dohet, A. Frantz, G. Geimer, M. Hellers, J. A. Massard, X. Mestdagh, R. Proess, N. Schneider & M. Pfeiffenschneider, 2017. Environmental impact assessment and black, watch and alert list classification after the ISEIA Protocol of invertebrates in Luxembourg. Bull. Soc. Nat. luxemb. 119: 63-70. [PDF 360 KB]
• Weber, D. (Hrsg.), 2014. Die Höhlenfauna Luxemburgs. Ferrantia 69. Musée national d’histoire naturelle Luxembourg, 408 pp. + CD-Rom (PDF 19 MB)
• Wikipedia contributors, 2019. Opilio canestrinii. Wikipedia, The Free Encyclopedia, 17 January 2019, 10:39 UTC, &lt;https://en.wikipedia.org/w/index.php?title=Opilio_canestrinii&amp;oldid=878854172&gt; [accessed 2020-03-05]

 Page content last updated on 2021-05-14. Last proofread by Caroline Grounds on 2019-12-10.

	Sheep tick	Status LU: absent
	Schofszeck	Status Eur.: established.
	n/a	RA: ISEIA: C0. Harmonia+: n/a.
	Schafzecke	Wikipedia: | Wikispecies:
	n/a	Back to the list of invertebrates

Important note

In 2015, six Dermacentor ticks were collected in SE-Luxembourg. Previously, these individuals had been identified as Dermacentor marginatus Leach, 1815. However, morphological re-investigation as well as DNA barcoding identified the specimens as Dermacentor reticulatus (Fabricius, 1794). Thus only D. reticulatus is known from Luxembourg (Weigand et al. 2020).

Brief description

Dermacentor marginatus Leach, 1815 is a species of tick of the family Ixodidae, which is a vector of several zoonotic diseases, which is why this tick is considered to pose particular eco-epidemiological and public health risks for humans (Wikipedia contributors 2019).

In Central Europe the sheep tick is the most common species of the genus. It lives here only in heat islands and on sunny slopes or dry grasslands. Mammals, including humans, are preferred as hosts. The animals of the genus transmit tularemia (Francisella tularensis), Rocky Mountain spotted fever (Rickettsia richettsii), Q-fever (Rickettsia burneti), bovine anaplasmosis, puppy-babesiosis (Babesia canis) (Source: http://www.ijon.de/zecken/system.html#zecke).

It is one of the species that could be favoured by global warming, which would allow it to move northwards, but also upwards (Wikipedia contributors 2019).

Status and distribution in Luxembourg

The tick Dermacentor marginatus Leach, 1815 is absent from Luxembourg.

Risk assessment

ISEIA protocol

Following the morphological re-investigation as well as DNA barcoding that identified all the specimens as Dermacentor reticulatus (Fabricius, 1794) (Weigand et al. 2020), Dermacentor marginatus is considered absent from Luxembourg and was reassessed on 29 October 2020 to C0 (2+1+2+1). Initial assessment: C1 (2+1+2+1) (Ries et al. 2017: 68).

Harmonia+ protocol

Not assessed yet.

Worldwide distribution

Bibliography

• Direction de la Santé, 2016. Attention aux tiques! Comment se protéger? Dépliant. Luxembourg. URL: https://sante.public.lu/fr/prevention/tiques/
• GBIF, 2020. Dermacentor marginatus in GBIF Secretariat (2019). GBIF Backbone Taxonomy. Checklist dataset https://doi.org/10.15468/39omei [accessed 2020-03-05]
• Ries, C., A. Arendt, C. Braunert, S. Christian, A. Dohet, A. Frantz, G. Geimer, M. Hellers, J. A. Massard, X. Mestdagh, R. Proess, N. Schneider & M. Pfeiffenschneider, 2017. Environmental impact assessment and black, watch and alert list classification after the ISEIA Protocol of invertebrates in Luxembourg. Bull. Soc. Nat. luxemb. 119: 63-70. [PDF 360 KB]
• Weigand, A., J. Teixeira & S. Christian, 2020. First record of Hyalomma marginatum sensu stricto C.L. Koch, 1844 and distribution of Dermacentor reticulatus (Fabricius, 1794) (Acari, Ixodidae) in Luxembourg. Bull. Soc. nat. luxemb. 122 : 253-263.
• Wikipedia contributors, 2019. Dermacentor marginatus. Wikipédia, l’encyclopédie libre. URL: https://fr.wikipedia.org/wiki/Dermacentor_marginatus [accessed 2020-03-05]

 Page content last updated on 2022-04-27. Last proofread by Caroline Grounds on 2019-12-10.

	Yellow sac spider1	Status LU: established. 1st record: 1988.
	Getëppelten Darfanger	Status Eur.: established.
	n/a	RA: ISEIA: C2. Harmonia+: n/a.
	Ammendornfinger	Wikipedia:   | Wikispecies: | wiki.arages.de
	Grote bermzakspin	Back to the list of invertebrates

Brief description

Cheiracanthium punctorium (Villers, 1789) lives in grasslands, preferring tall stalks. Being rather similar to several native Cheiracanthium species in its biology, C. punctorium should have no greater influence on the composition of native prey species in a colonised biotope. There may, however, be some competition with similar predators.

The species is known as quite aggressive when disturbed, especially pregnant or egg-protecting females. The chelicerae (fangs) of bigger individuals can easily penetrate through human skin and inject their venom. A bite of C. punctorium is painful and causes itching and swellings, and sometimes stronger reactions like vertigo, nausea, shivering, circulatory collapse and light fever. The symptoms usually decline after 24-30 hours. As the species leads a rather hidden lifestyle, the probability for bites should be rather small. There is a certain, but unfortunately sometimes overstated, media exposure of the species and its venomousness.

Status and distribution in Luxembourg

Cheiracanthium punctorium (Villers, 1789) was first documented in 1988 at the Aarnescht nature conservation reserve near Oberanven, municipality of Niederanven (Hermann 1998).

The species is rather thermophilous and therefore geographically restricted to the south of Luxembourg. Most observations have been made in dry grassland in the Moselle valley. Climate warming may favour further spread of the species.

Currently, 12 records of the species are accessible through the MNHNL-mdata portal; at the previously mentioned Aarnescht, and also at Roudebierg and Haard near Dudelange (MNHNL, iNaturalist & GBIF 2021).

Risk assessment

ISEIA protocol

C2 (2+2+2+1) (Ries et al. 2017: 68).

Harmonia+ protocol

Not assessed yet.

Distribution in Europe

Bibliography

• Arachnologische Gesellschaft, 2020. Atlas of the European Arachnids, accessed at https://atlas.arages.de [accessed 2020-03-05]
• Hermann, E., 1998. Die Spinnen (Araneae) ausgewählter Halbtrockenrasen im Osten Luxemburgs. Bull. Soc. Nat. luxemb. 99: 189-199. [PDF 165KB]
• MNHNL, 2000-. Cheiracanthium punctorium (Villers, 1789) in Recorder-Lux, database on the natural heritage of the Grand Duchy of Luxembourg. Musée national d’histoire naturelle, Luxembourg. URL: https://mdata.mnhn.lu [Accessed 2019-10-24]
• MNHNL, iNaturalist & GBIF, 2021. Cheiracanthium punctorium (Villers, 1789) in MNHNL-mdata, online portal combining species observation from Recorder-Lux, iNaturalist and GBIF. National Museum of Natural History, Luxembourg. URL: https://mdata.mnhn.lu [Accessed 2019-10-24]
• Ries, C., A. Arendt, C. Braunert, S. Christian, A. Dohet, A. Frantz, G. Geimer, M. Hellers, J. A. Massard, X. Mestdagh, R. Proess, N. Schneider & M. Pfeiffenschneider, 2017. Environmental impact assessment and black, watch and alert list classification after the ISEIA Protocol of invertebrates in Luxembourg. Bull. Soc. Nat. luxemb. 119: 63-70. [PDF 360 KB]
• Schneider, N., 2003. Auf Spurensuche. 22. Op der Aarnescht. Regulus 12/03: 22
• Thiel, M., 2016. Ein unheimlicher Zuwanderer. Eine südeuropäische Giftspinne zieht zunehmend nach Norden. Luxemburger Wort, 14.9.2016.

 Page content last updated on 2021-04-14. Last proofread by Caroline Grounds on 2019-12-05.

1. Cheiracanthium punctorium is one amongst several species commonly known as the yellow sac spider. Source: https://en.wikipedia.org/wiki/Cheiracanthium_punctorium.[↩]

	Wasp spider	Status LU: established. 1st record: 1906.
	Harespelspann	Status Eur.: established.
	Argiope frelon	RA: ISEIA: C3. Harmonia+: n/a
	Wespenspinne	Wikipedia: | Wikispecies: | CABI
	Wespspin	Back to the list of invertebrates

Brief description

Like many other members of its genus, Argiope bruennichi (Scopoli, 1772) shows striking yellow and black markings on its abdomen. The spider builds a spiral orb web at dawn or dusk, commonly in long grass a little above ground level (Wikipedia contributors 2019a).

Until around 50 years ago, the wasp spider was widespread in southern Europe, and very rarely present in central Europe. Since then the species has greatly enlarged and extended its area. Now it can be found in almost all European countries, as well as in some Asian and North African countries (Wikipedia contributors 2019b). The rapid spread of the species across Europe is generally thought to be facilitated by climate warming.

On sites where this strikingly marked spider occurs, the individual number can be very high. Argiope bruennichi populations might be a certain threat for rare species of their favourite prey and provoke changes in invertebrate communities of conquered sites. However the spider seems to integrate rather well in invaded countries, causing no great damage.

Status and distribution in Luxembourg

Argiope bruennichi (Scopoli, 1772) was first documented by Dr Ernest Feltgen on 24 August 1906 in Lintgen (Weinachter 1906; MNHNL 2000-).

Currently, 188 records of the wasp spider are accessible through the MNHNL-mdata portal (MNHNL, iNaturalist & GBIF 2019).

The species can be considered as widespread in the Grand Duchy.

Risk assessment

ISEIA protocol

C3 (2+2+2+2) (Ries et al. 2017: 68).

Harmonia+ protocol

Not assessed yet.

Worldwide distribution

Bibliography

• CABI, 2019. Argiope bruennichi. In: Invasive Species Compendium. Wallingford, UK: CAB International. URL: www.cabi.org/isc [accessed 2020-03-04]
• MNHNL, 2000-. Argiope bruennichi (Scopoli, 1772) in Recorder-Lux, database on the natural heritage of the Grand Duchy of Luxembourg. Musée national d’histoire naturelle, Luxembourg. URL: https://mdata.mnhn.lu [Accessed 2019-10-16]
• MNHNL, iNaturalist & GBIF, 2019. Argiope bruennichi (Scopoli, 1772) in MNHNL-mdata, online portal combining species observation from Recorder-Lux, iNaturalist and GBIF. National Museum of Natural History, Luxembourg. URL: https://mdata.mnhn.lu [Accessed 2019-10-16]
• Ries, C., A. Arendt, C. Braunert, S. Christian, A. Dohet, A. Frantz, G. Geimer, M. Hellers, J. A. Massard, X. Mestdagh, R. Proess, N. Schneider & M. Pfeiffenschneider, 2017. Environmental impact assessment and black, watch and alert list classification after the ISEIA Protocol of invertebrates in Luxembourg. Bull. Soc. Nat. luxemb. 119: 63-70. [PDF 360 KB]
• Weinachter, P., 1906. Comptes Rendus des Séances Fauna 16, 10: 215. [eluxembourgensia]
• Weiss, J., 1992. Die Ecke des Naturbeobachters. Regulus 3/92: 96-97.
• Wikipedia contributors, 2019a. Argiope bruennichi (Scopoli, 1772) in Wikipedia, The Free Encyclopedia. URL: https://en.wikipedia.org/w/index.php?title=Argiope_bruennichi&oldid=921986406 [accessed 24 October 2019]
• Wikipedia contributors, 2019b. Argiope bruennichi (Scopoli, 1772) in Wikipedia, Die freie Enzyklopädie. URL: https://de.wikipedia.org/w/index.php?title=Wespenspinne&oldid=192274595 [accessed 24 October 2019]

 Page content last updated on 2020-04-28. Last proofread by Caroline Grounds on 2019-12-04.

	Red swamp crayfish	Status LU: absent.
	Roude Suppekriibs	Status Eur.: established. IAS of EU concern (2016).
	Écrevisse de Louisiane	RA: ISEIA: A0, Alert List.
	Roter Amerikanischer Sumpfkrebs	Wikipedia: | Wikispecies: | CABI
	Rode rivierkreeft	Back to the list of invertebrates

Brief description

Procambarus clarkii (Girard, 1852) is a species of cambarid freshwater crayfish, native to northern Mexico, and southern and southeastern United States, but also introduced elsewhere, where it is often an invasive pest. P. clarkii is typically dark red, with long claws and head, small or no spines on the sides of its carapace just below the head, and rows of bright red bumps on the front and side of the first leg (Wikipedia contributors 2020).

The red swamp crayfish is most commonly found in warm fresh water, such as slowly flowing rivers, marshes, reservoirs, irrigation systems and rice paddies. It is considered to be the most ecologically plastic species in the order Decapoda, and is able to grow quickly even in only seasonally present water, being able to tolerate dry spells of up to four months. The red swamp crayfish grows quickly, and is capable of reaching weights in excess of 50 g, and sizes of 5.5–12 cm long. It is also able to tolerate slightly saline water, which is unusual for a crayfish. Additionally, P. clarkii are physiologically capable of tolerating relatively low dissolved oxygen concentrations. The average lifetime of Procambarus clarkii is five years (Wikipedia contributors 2020).

The burrowing activities of the red swamp crayfish can lead to damage to water courses and to crops, particularly rice, and its feeding can disrupt native ecosystems. It may out-compete the native crayfish species, and is a vector for the crayfish plague fungus Aphanomyces astaci, for crayfish virus vibriosis, and a number of worms parasitic on vertebrates. Their burrowing activities may also be a threat to civil infrastructure such as storm ponds and levees (Wikipedia contributors 2020).

Status and distribution in Luxembourg

There is no documented occurrence of Procambarus clarkii (Girard, 1852) in Luxembourg yet (MNHNL, iNaturalist & GBIF 2020).

Risk assessment

ISEIA protocol

A0 (3+3+3+3) = Alert List (Ries et al. 2017: 68).

Harmonia+ protocol

Not assessed yet.

Worldwide distribution

Bibliography

• CABI, 2019. Procambarus clarkii. In: Invasive Species Compendium. Wallingford, UK: CAB International. URL: www.cabi.org/isc [accessed 2020-03-13]
• MNHNL, iNaturalist & GBIF, 2020. Procambarus clarkii in MNHNL-mdata, online portal combining species observation from Recorder-Lux, iNaturalist and GBIF. National Museum of Natural History, Luxembourg. URL: https://mdata.mnhn.lu [accessed 2020-03-13]
• Ries, C., A. Arendt, C. Braunert, S. Christian, A. Dohet, A. Frantz, G. Geimer, M. Hellers, J. A. Massard, X. Mestdagh, R. Proess, N. Schneider & M. Pfeiffenschneider, 2017. Environmental impact assessment and black, watch and alert list classification after the ISEIA Protocol of invertebrates in Luxembourg. Bull. Soc. Nat. luxemb. 119: 63-70. [PDF 360 KB]
• Wikipedia contributors, 2020. Procambarus clarkii, Wikipedia, The Free Encyclopedia, 3 February 2020, 18:43 UTC, <https://en.wikipedia.org/w/index.php?title=Procambarus_clarkii&oldid=939000553> [accessed 2020-03-13]

 Page content last updated on 2021-05-14.